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Capricious Shiftings

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The Extermination of the American Bison, William Hornaday, 1889
The following is an excerpt from a book written by William Hornaday about the history of the American bison and its impending extinction.

Hornaday, William Temple. The Extermination of the American Bison. Washington: G.P.O., 1889. WorldCat [OCLC]. Web. 23 Mar. 2016.

It can be reasonably inferred that "capricious shiftings" in line 25 most nearly means

A

the subgroupings of bison within a herd.

B

the regional bison herds.

C

unexpected herd movements.

D

herd mating patterns.