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Select the best conjunctive adverb in the sentence below.

No one will dispute that instincts are of the highest importance to each animal;
Select Option thereforehoweveralthough
, there is no real difficulty, under changing conditions of life, in natural selection accumulating to any extent slight modifications of instinct which are in any way useful. Darwin, Charles Robert. The Origin of Species. New York: P.F. Collier & Son, 1909–14,
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