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What is the most likely meaning of the phrase "uncomfortably on all fours yet unheeding his discomfort" in the passage below?

Jack was bent double. He was down like a sprinter, his nose only a few inches from the humid earth. The tree trunks and the creepers that festooned them lost themselves in a green dusk thirty feet above him, and all about was the undergrowth. There was only the faintest indication of a trail here; a cracked twig and what might be the impression of one side of a hoof. He lowered his chin and stared at the traces as though he would force them to speak to him. Then dog-like, uncomfortably on all fours yet unheeding his discomfort, he stole forward five yards and stopped.

A

It reveals that Jack is so focused on tracking that he is unaware of his own pain.

B

It shows readers that Jack has become more animal than human.

C

It illustrates the incredible power the beast holds over Jack as he is beginning to look and act like him.

D

It implies that Jack is incredibly athletic and capable of superhuman feats in the jungle.

E

It suggests that while Jack wants to be a savage hunter, he has not yet fully made that transition.

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