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A number of structural features are listed below for hypothetical protein X. These features represent different levels of protein structure.

I. Aspartate at position 12 on subunit A interacts with lysine at position 98 on subunit B.
II. Cysteine at position 50 on subunit A forms a disulfide bond with cysteine at position 222 on subunit A.
III. At position 98, the peptide bond is planar.
IV. Positions 1 through 5 on subunit B contains methionine-glycine-alanine-arginine-leucine.

Which of the following describes a feature that relates to quaternary (${ 4 }^{ ° }$) structure?

A

I

B

II

C

III

D

IV

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